Research Interests

• Salmonid fish well-being in changing environment
• Ulcerative Dermal Necrosis (UDN)
• Biogeochemical Sensors (BGC Sensors)
• CO₂ system in the coastal zone
• Ecotoxicology

Current projects

• BENTOS - Multimodal Monitoring of the Baltic Sea Ecosystem. 2026-2030, co-investigator
• CLUSTER4FISH - Utilizing High-Performance Computing for the Study of Ulcerative Dermal Necrosis in Sea Trout. Stability AI. 2024-2026, project leader
• PROSPECTOR - do Permafrost-Released OrganicS amPlify ocEan aCidificaTiOn in the aRctic? National Science Center. 2020-2023, co-investigator
• WISLA - Transformations of the CO₂ system in the Vistula River Mouth. Polish National Science Centre. 2019-2021, project leader
• BONUS INTEGRAL - Integrated carboN and TracE Gas monitoRing for the bALtic sea, 2017-2020, co-investigator
• Modelling of the impact of the agricultural holdings and land-use structure on the quality of inland and coastal waters of the Baltic Sea set up on the example of the Municipality of Puck region - Integrated info-prediction Web Service WaterPUCK. The National Center for Research and Development, 2017-2020, co-investigator
• SALSA - Structure and functioning of the acid-base system in the Baltic Sea. Polish National Science Centre. 2016-2019, co-investigator

Recent publications

• Gozdowska, M., Stokowski, M., Sokolowska, E., Kulczykowska, E., 2026. Short communication: Skin melatonin and cortisol responses to water acidification and basification within the optimal pH range in three-spined sticklebacks. Comparative Biochemistry and Physiology, Part A 318, 112030, https://doi.org/10.1016/j.cbpa.2026.112030
• Uppuluri, K., Szwagierczak, D., Zaraska, K., Zachariasz, P., Stokowski, M., Synkiewicz-Musialska, B., Krzyœciak, P., 2025. pH Sensing Properties of Co3O4-RuO2-Based Electrodes and Their Application in Baltic Sea Water Quality Monitoring. Sensors 25, 1065, https://doi.org/10.3390/s25041065
• Aguado Gonzalo, F., Stokowski, M., Koziorowska-Makuch, K., Makuch, P., Beszczynska-Moller, A., Kuklinski, P., & Kulinski, K. (2024). Key processes controlling the variability of the summer marine CO₂ system in Fram Strait surface waters. Front. Mar. Sci., 11, https://doi.org/10.3389/fmars.2024.1464653
• Stokowski M., Sobiegraj W., Kulczykowska E. (2023). Potential role of climate change on the spread of salmonid skin condition: the biogeochemical hypothesis on ulcerative dermal necrosis on the Slupia River - Poland. Front. Mar. Sci., 10, https://doi.org/10.3389/fmars.2023.1104436
• Stokowski M., Winogradow A., Szymczycha B., Carstensen J., Kulinski K. (2021). The CO₂ System Dynamics in the Vicinity of the Vistula River Mouth (the southern Baltic Sea): A baseline investigation. Estuar. Coast. Shelf. Sci., 258, 107444, https://doi.org/10.1016/j.ecss.2021.107444
• Stokowski M., Makuch P., Rutkowski K., Wichorowski M., Kulinski K. (2021). A system for the determination of surface water pCO₂ in a highly variable environment, exemplified in the southern Baltic Sea. Oceanologia 63, 276-282, https://doi.org/10.1016/j.oceano.2021.01.001
• Stokowski M., Schneider B., Rehder G., Kulinski K. (2020). The CO₂ System Characteristics of the Oder River Estuary (the Baltic Sea). J. Mar. Syst., 211, 103418, https://doi.org/10.1016/j.jmarsys.2020.103418
• Szymczycha B., Klostowska Z., Kulinski K., Winogradow A., Jakacki J., Klusek Z., Grabowski M., Brodecka-Goluch A., Graca B., Stokowski M. (2018). Deep submarine groundwater discharge indicated by pore water chloride anomalies in the Gulf of Gdansk, southern Baltic Sea. E3S Web of Conferences, 54, 00035, https://doi.org/10.1051/e3sconf/20185400035
• Kulinski K., Schneider B., Szymczycha B., Stokowski M. (2017). Structure and functioning of the acid-base system in the Baltic Sea. Earth System Dynamics, 8, 1107-1120, https://doi.org/10.5194/esd-8-1107-2017
• Bialk-Bielinska A., Mulkiewicz E., Stokowski M., Stolte S., Stepnowski P. (2017). Acute aquatic toxicity assessment of six anti-cancer drugs and one metabolite using biotest battery - Biological effects and stability under test conditions. Chemosphere 189, 689-698, https://doi.org/10.1016/j.chemosphere.2017.08.174

Awards

• IAEA OA-ICC Travel Grant - Third Winter School on Ocean Acidification and Multiple Stressors, Monaco, 2025
• IOCCP Travel Grant, IOCCP & ICOS OTC Training course on a suite of biogeochemical sensors, Kristineberg, Sweden, 2023
• European Geosciences Union (EGU) Travel Grant, GO2NE SS2019: GO2NE Summer School, Xiamen, China, 2019
• Recognition for the poster presentation, International Sopot Youth Conference entitled "Where the World is Heading", Sopot, Poland, 2019
• IOCCP Travel Grant, GOA-ON Science Workshop in Hangzhou, China, 2019
• Award of the Director of the Institute of Oceanology PAN in Sopot for the activity in publishing scientific results, Sopot, Poland, 2017
• Professor Stanislaw Szymborski Award granted by the director of the Institute of Oceanology Polish Academy of Sciences for the best presentation in the field of natural sciences at the International Sopot Youth Conference, Sopot, Poland, 2017
• Third Best Poster Presentation at IV Young Researchers Conference on the occasion of World Water Day, Poznan, Poland, 2015

Patents and utility models

• Wejer, J., Dragan-Górska, A., Stokowski, M., Makuch, P., Markuszewski, P. (2025, patent pending). Multi-cell self-regenerating DC energy accumulator. Wieloogniwowy samoregenerujący się akumulator energii prądu stałego, [Urząd Patentowy Rzeczypospolitej Polskiej, application filed 14.01.2025, P.450950]
• Wejer, J., Dragan-Górska, A., Stokowski, M. (2024, utility model pending). Plate fuse with optical conduction monitor. Bezpiecznik płytkowy z optycznym monitorem przewodzenia, [Urząd Patentowy Rzeczypospolitej Polskiej, utility model application filed 20.06.2024, W.132204]
• Stokowski, M., Wejer, J., Żytko, A. (2022, patent pending). Device for electrochemical deacidification of water in water bodies based on an acid-base fuel cell. Urządzenie do elektrochemicznego odkwaszania wody w zbiornikach wodnych, stanowione paliwowym ogniwem kwasowo-zasadowym, [Urząd Patentowy Rzeczypospolitej Polskiej, application filed 08.07.2022, P.441683]
• Stokowski, M. (2021, patent pending). Container for calibration of an oxygen optode under anaerobic conditions. Pojemnik dla kalibracji optody tlenowej w warunkach środowiska beztlenowego, [WIPO ST 10/C PL433869]
• Wejer, J., Stokowski, M. (2019). Real-time wireless video transmission in aquatic environment. Układ bezprzewodowej transmisji w czasie rzeczywistym sygnału wizyjnego w środowisku wodnym, P.430971
• Stokowski, M., Wejer, J., Kuliński, K. (2018). System for monitoring the measuring instrument against the influence of unwanted moisture. Układ nadzorujący ustrój instrumentu pomiarowego i zabezpieczający go przed wpływem niepożądanej wilgoci, P.425618

Trainings and courses

• IAEA OA-ICC - Third Winter School on Ocean Acidification and Multiple Stressors, Monaco, 2025
• IOCCP & ICOS OTC Training course on a suite of biogeochemical sensors, Sweden, 2023
• EdinburghX: Climate Change: Carbon Capture and Storage Training, online, 2023
• Training on planning procedures and experiments and carrying them out; performing the procedures; killing animals used in procedures, The Polish Laboratory Animal Science Association (PolLASA), online, 2022
• Training on supervising the welfare and care of animals kept at the facility; caring for animals kept at the facility, The Polish Laboratory Animal Science Association (PolLASA), online, 2022
• On a scale of 0-14, how familiar are you with ocean acidification pHacts?! course, NOAA Ocean Acidification Program, online, 2020
• International Summer School of the Global Ocean Oxygen Network (GO2NE), China, 2019
• Workshop on Underway CO₂ data and metadata quality control procedures, Institute of Oceanology Polish Academy of Science (IO PAN), Poland, 2019
• 4th International Summer School on Climate of the Baltic Sea Region, Sweden, 2018
• Course Safe handling of sealed sources of ionizing radiation, Poland, 2013
• Course Accuracy, precision in mass measurements, Poland, 2013
• Course New analytical methods, Poland, 2013
• Summer course on Ichthyology of the sea, Professor Krzysztof Skora Marine Station of the Institute of Oceanography, Poland, 2013

Conference Presentations

1. M. Stokowski. (2026). Could Climate-Driven Changes in Estuarine Water Chemistry Contribute to Ulcerative Dermal Necrosis in Salmonids? 13th Societas Humboldtiana Polonorum Congress Gdansk, 2026
2. M. Stokowski, E. Kulczykowska, M. Gozdowska, P. Debowski, M. Wichorowski, M. Hoeft, K. Gierlowski, R. Bernas, A. Zytko, D. Rak. (2025). Investigating Climate Change-Driven Biogeochemical Variability in Estuaries as a Potential Component of UDN Aetiology in Salmonid Fish, The HealthyRiver Project. Baltic Sea Science Congress, Sopot, Poland, oral presentation
3. M. Stokowski. (2024). Fascynujacy swiat ryb lososiowatych: biologia, ekologia i ochrona, (Nie)zrownowazony Baltyk webinar series, oral presentation
4. M. Stokowski. (2023). Salmonid Fish - Fascinating Wanderers. Conference 2023 entitled Where the World is Heading, Online. OPENING SPEECH.
5. Palka, K. Koziorowska-Makuch, F. Aguado Gonzalo, P. Makuch, M. Stokowski, S. R. Saghravani, B. Szymczycha, A. Winogradow, K. Kulinski. (2022). Charakterystyka rozpuszczonej materii organicznej uwalnianej z wieloletniej zmarzliny (Spitsbergen). III Konferencja Naukowa Polskich Badaczy Morza, Gdynia, Poland, poster
6. F. Aguado Gonzalo, M. Stokowski, K. Koziorowska-Makuch, P. Kuklinski, P. Makuch, I. Palka, B. Szymczycha, K. Kulinski. (2022). Biological and physical processes affecting pH in the eastern part of the Fram Strait area. III Konferencja Naukowa Polskich Badaczy Morza, Gdynia, Poland, poster.
7. F. Aguado Gonzalo, M. Stokowski, K. Koziorowska-Makuch, P. Kuklinski, P. Makuch, I. Palka, B. Szymczycha, K. Kulinski. (2022). The importance of sea ice in the processes controlling pH in the Fram Strait Area. Ocean Biogeochemistry, Gordon Research Conference, Biogeochemical Processes Across Space and Time, Barcelona, Spain, poster
8. M. Stokowski, W. Sobiegraj, M. Gozdowska, A. Winogradow, E. Kulczykowska. (2022). Czy zmiany klimatyczne moga miec zwiazek z wrzodziejaca martwica skory (UDN) ryb lososiowatych? Problemy Zdrowotne Tarlakow Troci i Lososia - Ocena Zagrozenia, Poznan, Poland, oral presentation
9. M. Stokowski, W. Sobiegraj, M. Gozdowska, A. Winogradow, E. Kulczykowska. (2022). Czy zmiany klimatyczne moga miec zwiazek z wrzodziejaca martwica skory (UDN) ryb lososiowatych? III Konferencja Naukowa Polskich Badaczy Morza, Gdynia, Poland, oral presentation
10. M. Stokowski, B. Schneider, G. Rehder, A. Winogradow, B. Szymczycha, J. Carstensen, K. Kulinski. (2022). The marine CO2 system in the estuaries of the Vistula and Oder Rivers - Structure, variability and potential implications for the Baltic Sea. 4th Baltic Earth Conference, Jastarnia, Poland, oral presentation
11. M. Stokowski, A. Winogradow, B. Szymczycha, J. Carstensen, K. Kulinski. (2021). The CO2 system dynamics in the vicinity of the Vistula River mouth (the southern Baltic Sea): a baseline investigation. Baltic Sea Science Congress, Denmark, oral presentation
12. M. Stokowski, A. Winogradow, B. Szymczycha, K. Kulinski. (2020). The CO2 system variability in the vicinity of the Vistula River mouth. Earth System Changes and Baltic Sea Coasts, Online, poster
13. M. Stokowski, A. Winogradow, B. Szymczycha, K. Kulinski. (2020). The CO2 System Variability in the Vicinity of the Vistula River Mouth. Goldschmidt Conference, Online, poster
14. M. Stokowski, A. Winogradow, B. Szymczycha, K. Kulinski. (2020). Transformacje systemu weglanowego w ujsciu Wisly. Chemia, Geochemia i Ochrona Srodowiska Morskiego, Online, poster
15. M. Stokowski, K. Kulinski. (2019). Pomiary pCO2 w wodzie. XXIV Konferencja: Nowoczesne Metody Instrumentalne w Analizie Sladowej, Lodz, Poland, oral presentation
16. M. Stokowski, K. Kulinski. (2019). Charakterystyka i przeksztalcenia systemu weglanowego w ujsciach Wisly i Odry. II Konferencja Naukowa Polskich Badaczy Morza, Gdynia, Poland, oral presentation
17. M. Stokowski, A. Winogradow, K. Kulinski. (2019). The characteristics of the carbonate system in the estuary of the Vistula River (Poland). Baltic Sea Science Congress, Stockholm, Sweden, oral presentation
18. M. Stokowski, K. Kulinski. (2019). Transformations of the carbonate system in the estuary of the Odra and Vistula rivers. INTERNATIONAL SOPOT YOUTH CONFERENCE, Sopot, Poland, oral presentation
19. M. Stokowski, K. Kulinski, B. Schneider, G. Rehder, A. Winogradow, M. Lengier. (2019). The dynamics of the CO2 system in the coastal zone of the Southern Baltic Sea. Global Ocean Oxygen Network GO2NE Summer School, Xiamen, China, poster
20. M. Stokowski, B. Schneider, G. Rehder, K. Kulinski. (2019). Transformations of the carbonate system in the estuary of the Odra and Vistula Rivers (The Baltic Sea). 4th GOA-ON International Workshop, Hangzhou, China, poster
21. M. Stokowski, B. Schneider, K. Kulinski. (2018). Drivers of the CO2 system in the coastal zone of the Baltic Sea. Baltic Earth Workshop on Multiple drivers for Earth system changes in the Baltic Sea region, Tallinn, Estonia, oral presentation
22. M. Stokowski, B. Schneider, K. Kulinski. (2018). Drivers of the CO2 system in the coastal zone of the Baltic Sea. Baltic Earth Workshop on Multiple drivers for Earth system changes in the Baltic Sea region, Tallinn, Estonia, poster
23. M. Stokowski, D. Ren, O. Vinogradova, E. Mikheeva, L. Naumov. (2018). Summer 2018: a trend or an event? 4th International Baltic Earth Summer School on Climate of the Baltic Sea region, Asko, Sweden, oral presentation
24. M. Stokowski. (2018). My PhD thesis work. 4th International Baltic Earth Summer School on Climate of the Baltic Sea region, Asko, Sweden, oral presentation
25. Szymczycha, Z. Klostowska, K. Kulinski, A. Winogradow, J. Jakacki, Z. Klusek, M. Grabowski, A. Brodecka-Goluch, B. Graca, M. Stokowski. (2018). Deep submarine groundwater discharge indicated by pore water chloride anomalies in the Gulf of Gdansk, southern Baltic Sea. 25th Salt Water Intrusion Meeting, Gdansk, Poland, oral presentation
26. K. Kulinski, M. Stokowski, B. Szymczycha, K. Hammer, K. Koziorowska, A. Winogradow, M. Lengier, Z. Klostowska, B. Schneider. (2018). The acid-base system of the Baltic Sea. 2nd Baltic Earth Conference: The Baltic Sea Region in Transition, Helsingor, Denmark, oral presentation
27. Szymczycha, Z. Klostowska, K. Kulinski, A. Winogradow, J. Jakacki, Z. Klusek, M. Grabowski, A. Brodecka-Goluch, B. Graca, M. Stokowski, K. Koziorowska, D. Rak. (2018). Deep submarine groundwater discharge indicated by pore water chloride anomalies in the Gulf of Gdansk, southern Baltic Sea. 2nd Baltic Earth Conference: The Baltic Sea Region in Transition, Helsingor, Denmark, oral presentation
28. Winogradow, B. Szymczycha, K. Koziorowska, A. Mackiewicz, M. Stokowski, K. Kulinski, J. Pempkowiak. (2018). Stale izotopy wegla w badaniach pochodzenia materii organicznej w Morzu Baltyckim. XIII Konferencja: Chemia, Geochemia i Ochrona Srodowiska Morskiego, Sopot, Poland, oral presentation
29. Winogradow, B. Szymczycha, K. Koziorowska, A. Mackiewicz, M. Stokowski, K. Kulinski, J. Pempkowiak. (2018). Osady Morza Baltyckiego jako zrodlo C, N, P. XIII Konferencja: Chemia, Geochemia i Ochrona Srodowiska Morskiego, Sopot, Poland, oral presentation
30. M. Stokowski, B. Schneider, K. Kulinski. (2018). Charakterystyka systemu weglanowego w ujsciu Odry. XIII Konferencja: Chemia, Geochemia i Ochrona Srodowiska Morskiego, Sopot, Poland, oral presentation
31. K. Kulinski, B. Szymczycha, A. Winogradow, M. Stokowski, K. Koziorowska, J. Pempkowiak. (2017). Struktura i funkcjonowanie systemu kwasowo-zasadowego w Morzu Baltyckim. I Konferencja Naukowa Polskich Badaczy Morza, Sopot, Poland, oral presentation
32. Winogradow, B. Szymczycha, K. Koziorowska, M. Stokowski, K. Kulinski, J. Pempkowiak. (2017). Stabilne izotopy wegla jako wskazniki pochodzenia materii organicznej w Morzu Baltyckim. I Konferencja Naukowa Polskich Badaczy Morza, Sopot, Poland, oral presentation
33. M. Stokowski, B. Schneider, J. Muller, G. Rehder, K. Kulinski. (2017). Charakterystyka systemu weglanowego w estuarium rzeki Odry. I Konferencja Naukowa Polskich Badaczy Morza: Stan i trendy zmian w morzach i oceanach, Sopot, Poland, poster
34. M. Stokowski, B. Schneider, J. Muller, G. Rehder, K. Kulinski. (2017). The characteristics of the carbonate system in the Odra river estuary (Poland). The 11th Baltic Sea Science Congress: Living along gradients: past, present, future, Rostock, Germany, poster
35. K. Kulinski, B. Schneider, B. Szymczycha, K. Hammer, A. Winogradow, M. Stokowski, K. Koziorowska. (2017). Structure and functioning of the acid-base system in the Baltic Sea. The 11th Baltic Sea Science Congress: Living along gradients: past, present, future, Rostock, Germany, oral presentation
36. Bialk-Bielinska, E. Mulkiewicz, M. Stokowski, L. Grabarczyk, M. Pazda, P. Stepnowski. (2017). Chromatografia cieczowa jako narzedzie w badaniach stabilnosci i biodostepnosci wybranych lekow przeciwnowotworowych o charakterze jonowym i ich produktow transformacji podczas testow w ocenie ich ryzyka ekotoksykologicznego wobec rzesy wodnej L. minor. V Konferencja Naukowa: Monitoring i analiza wody. Chromatograficzne metody oznaczania substancji o charakterze jonowym, Lysomice k. Torunia, Poland, poster
37. Bialk-Bielinska, K. Mioduszewska, J. Dolzonek, M. Tonski, M. Stokowski, E. Mulkiewicz, P. Stepnowski. (2017). Liquid chromatography in the modeling studies on the mobility, hydrolytic stability and ecotoxicity of selected anticancer drugs and their transformation products in the environment. XI Polska Konferencja Chromatograficzna and 24th International Symposium on Electro- and Liquid Phase-Separation Techniques, Sopot, Poland, poster
38. Bialk-Bielinska, K. Mioduszewska, J. Dolzonek, M. Tonski, M. Stokowski, E. Mulkiewicz, P. Stepnowski. (2017). Assessment of mobility, hydrolytic stability and ecotoxicity of anticancer drugs and their transformation products in the environment. VIII Miedzynarodowa Konferencja Naukowa: Toxic substances in the environment, Krakow, Poland, oral presentation
39. K. Kulinski, B. Schneider, B. Szymczycha, K. Hammer, A. Winogradow, M. Stokowski, K. Koziorowska. (2017). Structure and functioning of the acid-base system in the Baltic Sea. The 11th Baltic Sea Science Congress, Rostock, Germany. ORAL PRESENTATION.
40. K. Kulinski, B. Szymczycha, A. Winogradow, M. Stokowski, K. Koziorowska, J. Pempkowiak. (2017). Struktura i funkcjonowanie systemu kwasowo-zasadowego w Morzu Baltyckim. I Konferencja Naukowa Polskich Badaczy Morza, Sopot, Poland. ORAL PRESENTATION.
41. Winogradow, B. Szymczycha, K. Koziorowska, M. Stokowski, K. Kulinski, J. Pempkowiak. (2017). Stabilne izotopy wegla jako wskazniki pochodzenia materii organicznej w Morzu Baltyckim. Ograniczenia zwi¹zane z obszarem badan, sezonowosci¹ oraz glebokosci¹. I Konferencja Naukowa Polskich Badaczy Morza, Sopot, Poland, oral presentation
42. M. Stokowski, B. Schneider, J. Muller, G. Rehder, K. Kulinski. (2017). Charakterystyka systemu weglanowego w estuarium rzeki Odry. I Konferencja Naukowa Polskich Badaczy Morza, Sopot, Poland. POSTER.
43. M. Stokowski, B. Schneider, J. Muller, G. Rehder, K. Kulinski. (2017). The characteristics of the carbonate system in the Odra river estuary (Poland). International Sopot Youth Conference, Sopot, Poland, oral presentation
44. M. Stokowski, K. Kulinski. (2016). Porownanie dwoch metod pomiaru pCO2 w wodzie morskiej opartych na pelnej oraz czesciowej rownowadze fazy cieklej i gazowej. XII Konferencja: Chemia, Geochemia i Ochrona Srodowiska Morskiego, Sopot, Poland, oral presentation
45. K. Kulinski, B. Schneider, B. Szymczycha, K. Hammer, A. Winogradow, M. Stokowski, K. Koziorowska. (2016). Peculiarities of the Baltic Sea acid-base system. 1st Baltic Earth Conference, Nida, Lithuania, oral presentation
46. M. Stokowski, A. Puckowski, S. Stolte, P. Stepnowski, A. Bialk-Bielinska. (2015). Aquatic toxicity of six cytostatic drugs commonly used in treatment of human cancer. Swiatowe Dni Wody, Poznan, Poland, poster